Carly Cooper is 30, single and after coming close to saying ‘I do’ to six different men, she’s wondering if she accidentally said ‘goodbye' to Mr Right. But there is a problem. Her ex-boyfriends are scattered all over the world and Carly lives in 1999: an era before Facebook, Google, smartphones, 4G and Broadband, when it was impossible to track people down with a few clicks of a mouse. On a mission to discover if she walked away from her 'happy ever after', Carly quits her job, her flat, her whole life and sets off on a quest to track down all the men she has ever loved.

Tress Walker waved her perfect husband Max off to work, with no idea that she was about to go into labour with their first child. And completely unaware that when she tried to track Max down, he wouldn’t be where he was supposed to be. At the same time, Max’s best friend Noah Clark said goodbye to his wife, Anya, blissfully oblivious that he would soon discover the woman he adored had been lying to him for years. And living alongside the two couples, their recently widowed friend, Nancy Jenkins, is getting ready to meet Eddie, her first true love at a school reunion.

After the loss of her mother and her own battle with breast cancer, Joanna Teale returns to her graduate research on nesting birds in rural Illinois, determined to prove that her recent hardships have not broken her. She throws herself into her work from dusk to dawn, until her solitary routine is disrupted by the appearance of a mysterious child who shows up at her cabin barefoot and covered in bruises. The girl calls herself Ursa, and she claims to have been sent from the stars to witness five miracles.

Colm O’Flynn was loved by his close circle of family and friends, however his death came too soon for everyone to make peace with their past. Shauna, his second wife, adored him. But one night she broke their marriage vows and didn’t get time to ask Colm’s forgiveness. Jess was the first Mrs O’Flynn. Her heart is set on someone new, but will the last one-night stand she shared with Colm come back to haunt her? Colm’s best friend, Dan, is recently divorced. Can he take a second shot at happiness if it means betraying the one person who always had his back?

Twenty years ago, Carly Cooper went on an epic quest to track down all the men she’d ever loved and lost in the hope that one of them was her Mr Right. Now, two decades and two teenage sons later, she thinks she might have got it all wrong. As the years have passed, lots of things have changed, leaving Carly asking, ‘What now?’ With a divorce and an empty nest on the horizon, Carly sets off once more to Los Angeles with her band of trusty girlfriends, to find the carefree, wild and adventurous Carly Cooper that she used to be.